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Artwork: Geoffrey Cottenceau and Romain Rousset, Flamme, 2009 In 1976, evolutionary biologist Richard Dawkins wrote in The Selfish Gene, “If you wish, as I do, to build a society in which individuals cooperate generously and unselfishly towards a common good, you can expect little help from biological nature. Let us try to teach generosity and altruism, because we are born selfish.” By 2006, the tide had started to turn. Harvard University mathematical biologist Martin Nowak could declare, in an overview of the evolution of cooperation in Science magazine, “Perhaps the most remarkable aspect of evolution is its ability to generate cooperation in a competitive world. Thus, we might add ‘natural cooperation’ as a third fundamental principle of evolution beside mutation and natural selection.”

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The Unselfish Gene

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